On attached pic highlighted is download queue tree for traffic with connection mark large. Window beneath the queue tree windows is connections. As shown there are no large connections listed yet queue is passing >800kbps of traffic.
On the other hand Upload side lists large connections just fine (second pic). Is this a bug?
A stateful connection for TCP traffic shows as the src IP address the IP address of the host that sent the initial SYN. In your case it looks like 18x.x.x.x/8 were the traffic initiators for all connections.
Mangle rules see packets, which can have the source address of either endpoint as they are unidirectional. Connection tracking sees what for TCP is called virtual circuits, which are bidirectional and have the source address of the endpoint that initiated the connection.
Mangle rules can access the state of a connection via the services provided by the connection tracking facility and use the state for a rule decision, but they inherently deal with single packets.
I still think it’s not right that dst-src and src-dst are not listed under connections tab. In addition how can one explain pic below: config same as above pics - traffic based on large connection mark yet there are no connections with large mark listed under connections at all: