RouterOS L2TP client interface question

Hello.
We’ve tried to connect hAP AC with LTE USB modem as a L2TP client to our VPN L2 overlay based on SoftEther server in public cloud.
We have 10.0.0.0/15 (production segment) and 192.168.88.0/24 (wifi segment) networks behind hAP. VPN overlay network is 192.168..168.0/24.
There is no problem with L2TP connectivity - it works well. So when we try to ping hosts in VPN network from production or wifi segments interfaces we have no success, it works only with masquerading on l2tp virtual interface.
Could you help with explanation on can we and how we can use hAP in routed mode (without NAT) as a gateway for production and wifi internal networks to L2TP VPN network?
Appreciate your help.

Here is routs for info:

[admin@MikroTik] > ip route print
Flags: X - disabled, A - active, D - dynamic, C - connect, S - static, r - rip, b - bgp, o - ospf, m - mme, B - blackhole, U - unreachable, P - prohibit
 #      DST-ADDRESS        PREF-SRC        GATEWAY            DISTANCE
 0 ADS  0.0.0.0/0                          192.168.0.1               0
 1  Do  0.0.0.0/0                          172.16.0.1              110
 2 ADC  1.0.0.1/32         192.168.168.12  l2tp-v2.xx.xx.xx          0
 3 ADo  10.0.1.0/24                        172.16.0.1              110
 4 ADo  10.0.2.0/23                        172.16.0.1              110
 5 ADo  10.0.9.0/28                        172.16.0.1              110
 6 ADo  10.0.9.16/28                       172.16.0.1              110
 7 ADo  10.0.9.32/28                       172.16.0.1              110
 8 ADo  10.0.9.48/28                       172.16.0.1              110
 9 ADo  10.0.9.64/28                       172.16.0.1              110
10 ADo  10.0.9.80/28                       172.16.0.1              110
11 ADo  10.0.9.96/28                       172.16.0.1              110
12 ADo  10.0.64.0/24                       172.16.0.1              110
13 ADo  10.0.65.0/24                       172.16.0.1              110
14 ADo  10.0.72.0/28                       172.16.0.1              110
15 ADo  10.0.72.16/28                      172.16.0.1              110
16 ADo  10.0.72.32/28                      172.16.0.1              110
17 ADo  10.0.72.48/28                      172.16.0.1              110
18 ADo  10.0.72.64/28                      172.16.0.1              110
19 ADo  10.0.72.80/28                      172.16.0.1              110
20 ADo  10.0.80.0/26                       172.16.0.1              110
21 ADo  10.0.96.0/27                       172.16.0.1              110
22 ADo  10.0.96.32/28                      172.16.0.1              110
23 ADo  10.0.96.48/28                      172.16.0.1              110
24 ADo  10.0.96.64/32                      172.16.0.1              110
25 ADo  10.0.97.0/30                       172.16.0.1              110
26 ADo  10.0.97.4/30                       172.16.0.1              110
27 ADo  10.0.99.0/24                       172.16.0.1              110
28 ADo  10.1.1.0/24                        172.16.0.1              110
29 ADo  10.1.2.0/23                        172.16.0.1              110
30 ADo  10.1.9.0/28                        172.16.0.1              110
31 ADo  10.1.9.16/28                       172.16.0.1              110
32 ADo  10.1.9.32/28                       172.16.0.1              110
33 ADo  10.1.9.48/28                       172.16.0.1              110
34 ADo  10.1.9.64/28                       172.16.0.1              110
35 ADo  10.1.9.80/28                       172.16.0.1              110
36 ADo  10.1.96.32/28                      172.16.0.1              110
37 ADo  10.1.96.64/32                      172.16.0.1              110
38 ADC  172.16.0.0/16      172.16.0.2      ether1                    0
39 ADC  192.168.0.0/24     192.168.0.104   lte1                      0
40 ADC  192.168.88.0/24    192.168.88.1    bridge                    0
41 A S  192.168.168.0/24                   l2tp-v2.xx.xx.xx          1