I am constructing my first WDS network, linking four total routers in a repeater-like chain, as shown in the following illustration:
Station-WDS — AP-bridge — AP-bridge — Station-WDS

I have set both interior APs as AP-bridge, and enabled dynamic mesh WDS on them, pointing to the local bridge interface (bridging wlan1 and ether ports). Similarly, the two end stations are set to Station-WDS, with same/similar settings. Bridge interfaces on all routers are on the same subnet.
Dynamic WDS links do get established automatically, but I cannot ping one bridge interface from one router to the other reliably. However, when I add a specific route for the next gateway (redundant with the subnet route) on a specific router (as shown in this review of the first link), it will work.
Attached are screen snapshots of the first link in this network, showing when I get no ping connection, and when I do after adding the gateway-specific route. We are looking at the first link in the chain.
Here, in this image of the first router, you can see packets leaving the router R1 for the ping target R2 But nothing coming back. Note the static route to the R2 bridge interface is disabled, and the dynamic route to the .88. subnet is active. On R2, no ping packets arrive from R1 (as seen via Firewall Log rules).
In image of the second router R2 pinging R1, you see a similar result - no ping connection.

But, when you look at R1 while being pinged by R2, you see that ping packets arrive and leave with the destination of R2 - but obviously don’t get there!

Here, I have enabled the ‘redundant’ gateway specif route on R1 to the bridge interface on R2 (tried the same trick on R2, but no effect), and you can see that now ping works from both directions. Here’s the view from R1:

… and here pinging from R2:

As I understand the manual and the illustrations of WDS networks there, I should be able to ping from any router to another to bridge interfaces on this common subnet (.88.), without having to ‘monkey’ with the routes. Clearly, something is amiss, and I have little confidence in this network until I get this resolved.
What do I have wrong here? The other links in this WDS repeater chain have similar issues…
Thanks in advance for the input…