I guess that what originated the misunderstanding is that the Fritzbox uses something called UPnP AV (Universal Plug and Play Audio/Video, which is NOT "plain" UPnP, but rather an extension of it): https://en.wikipedia.org/wiki/Universal_Plug_and_Play#AV_standards
It seems like some special configuration is needed to allow the packets/discovery/whatever to pass through:
though I couldn't find anything confirmed about this stuff.
There is a (as usual telegraphic) article on Mikrotik help about Ros being able to do the same (DLNA) media server but nothing about the needed settings on other devices on the network:
The thing revolves around SSDP, so maybe, just maybe, there could be something useful here: